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        "year": "2021",
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        "leaf_water_content_per_saturated_mass",
        "osmotic_potential_at_full_turgor",
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        "leaf_dark_respiration_per_area",
        "leaf_dark_respiration_per_dry_mass",
        "leaf_intercellular_co2_concentration_at_amax",
        "leaf_intercellular_co2_concentration_at_asat",
        "leaf_intercellular_co2_concentration_to_atmospheric_co2_concentration_ratio",
        "leaf_photosynthetic_nitrogen_use_efficiency_saturated",
        "leaf_photosynthetic_phosphorus_use_efficiency_saturated",
        "leaf_photosynthetic_rate_per_area_maximum",
        "leaf_photosynthetic_rate_per_area_saturated",
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        "leaf_photosynthetic_rate_per_dry_mass_saturated",
        "leaf_transpiration_per_area_at_asat",
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        "leaf_transpiration_per_area_at_amax",
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        "stem_water_potential_50percent_lost_conductivity",
        "stem_water_potential_88percent_lost_conductivity",
        "leaf_dry_mass",
        "recruitment_time",
        "leaf_lamina_division",
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        "huber_value",
        "leaf_specific_hydraulic_conductivity",
        "leaf_conductance_surface_water",
        "leaf_foliar_water_uptake_rate",
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        "leaf_cuticular_conductance",
        "leaf_fresh_mass",
        "water_potential_midday",
        "water_potential_predawn",
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        "leaf_water_content_per_dry_mass",
        "bark_density",
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        "stem_saturated_water_content_per_dry_mass",
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        "leaf_cn_ratio",
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        "leaf_delta15n",
        "wood_c_per_dry_mass",
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        "fire_time_from_fire_to_flowering",
        "fire_time_from_fire_to_flowering_decline",
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        "dispersal_unit",
        "dispersers",
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        "plant_flood_regime_classification",
        "vegetative_reproduction_ability",
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        "leaf_photosynthesis_vcmax_per_area",
        "lifespan",
        "resprouting_capacity_juvenile",
        "seedling_establishment_conditions",
        "life_form",
        "leaf_photosynthesis_jmax_per_area",
        "diaspore_fleshiness",
        "fruit_fleshiness",
        "leaf_photosynthetic_rate_per_area_ambient",
        "leaf_stomatal_conductance_per_area_ambient",
        "leaf_transpiration_per_area_ambient",
        "stem_hydraulic_conductivity",
        "hydraulic_safety_margin_50",
        "fire_time_from_fire_to_fruiting",
        "fruiting_time",
        "plant_diameter_breast_height",
        "serotiny",
        "leaf_water_potential_12percent_lost_conductivity",
        "root_water_potential_12percent_lost_conductivity",
        "root_water_potential_50percent_lost_conductivity",
        "root_water_potential_88percent_lost_conductivity",
        "root_vascular_anatomy",
        "leaf_light_respiration_per_area",
        "leaf_senesced_n_per_dry_mass",
        "leaf_senesced_p_per_dry_mass",
        "wood_p_per_dry_mass",
        "cell_cross_sectional_area",
        "leaf_cuticle_thickness_abaxial",
        "leaf_cuticle_thickness_adaxial",
        "leaf_epidermis_thickness_abaxial",
        "leaf_epidermis_thickness_adaxial",
        "leaf_palisade_tissue_thickness_abaxial",
        "leaf_palisade_tissue_thickness_adaxial",
        "leaf_spongy_mesophyll_thickness",
        "nitrogen_fixing",
        "leaf_hypodermis_thickness_adaxial",
        "leaf_hypodermis_thickness_abaxial",
        "leaf_inclination_angle",
        "leaf_stomatal_resistance_ambient",
        "leaf_reflectance",
        "leaf_reflectance_near_infrared",
        "leaf_water_content_per_fresh_mass",
        "leaf_lifespan",
        "seed_p_per_seed_dry_mass",
        "leaf_b_per_dry_mass",
        "leaf_ca_per_dry_mass",
        "leaf_cu_per_dry_mass",
        "leaf_fe_per_dry_mass",
        "leaf_k_per_dry_mass",
        "leaf_mg_per_dry_mass",
        "leaf_mn_per_dry_mass",
        "leaf_ni_per_dry_mass",
        "leaf_s_per_dry_mass",
        "leaf_zn_per_dry_mass",
        "leaf_mo_per_dry_mass",
        "leaf_si_per_dry_mass",
        "leaf_ash_per_dry_mass",
        "leaf_phenol_per_dry_mass",
        "leaf_tannin_per_dry_mass",
        "leaf_work_to_shear",
        "leaf_work_to_shear_adjusted",
        "leaf_area_ratio",
        "integrated_plant_transpiration",
        "root_shoot_ratio",
        "leaf_soluble_starch_per_mass",
        "leaf_soluble_sugars_per_mass",
        "root_soluble_starch_per_mass",
        "root_soluble_sugars_per_mass",
        "stem_soluble_starch_per_mass",
        "stem_soluble_sugars_per_mass",
        "leaf_stomatal_distribution",
        "leaf_work_to_punch",
        "leaf_work_to_punch_adjusted",
        "leaf_work_to_tear_adjusted",
        "atmospheric_co2_concentration",
        "leaf_chloroplast_co2_concentration",
        "leaf_photosynthesis_jmax_per_mass",
        "leaf_photosynthesis_vcmax_per_mass",
        "osmotic_potential",
        "root_mass_fraction",
        "stem_mass_fraction",
        "leaflet_dry_mass",
        "leaflet_area",
        "branch_mass_fraction",
        "plant_succulence",
        "leaf_mass_fraction",
        "leaf_relative_water_content_predawn",
        "leaf_water_use_efficiency_integrated",
        "leaf_stomatal_density_abaxial",
        "leaf_stomatal_density_adaxial",
        "leaf_stomatal_density_average",
        "resprouting_capacity_proportion_individuals",
        "root_structure",
        "root_system_classification",
        "root_specific_root_length",
        "genome_size",
        "root_wood_density",
        "leaf_chlorophyll_a_b_ratio",
        "leaf_chlorophyll_per_area",
        "leaf_soluble_protein_per_area",
        "leaf_soluble_starch_per_area",
        "leaf_soluble_sugars_per_area",
        "fruit_p_per_dry_mass",
        "competitive_stratum",
        "fire_time_from_fire_to_50_percent_fruiting",
        "leaf_na_per_dry_mass",
        "fire_flame_duration",
        "fire_smoulder_duration",
        "fire_time_to_ignition",
        "fire_total_burn_duration",
        "leaf_density",
        "leaf_lignin_per_dry_mass",
        "bark_al_per_dry_mass",
        "bark_b_per_dry_mass",
        "bark_c_per_dry_mass",
        "bark_ca_per_dry_mass",
        "bark_cu_per_dry_mass",
        "bark_fe_per_dry_mass",
        "bark_k_per_dry_mass",
        "bark_mg_per_dry_mass",
        "bark_mn_per_dry_mass",
        "bark_n_per_dry_mass",
        "bark_na_per_dry_mass",
        "bark_p_per_dry_mass",
        "bark_s_per_dry_mass",
        "bark_zn_per_dry_mass",
        "bark_ash_per_dry_mass",
        "bark_cellulose_per_dry_mass",
        "bark_lignin_per_dry_mass",
        "bark_tannin_per_dry_mass",
        "bark_water_content_per_dry_mass",
        "leaf_cellulose_per_dry_mass",
        "leaf_senesced_al_per_dry_mass",
        "leaf_senesced_b_per_dry_mass",
        "leaf_senesced_c_per_dry_mass",
        "leaf_senesced_ca_per_dry_mass",
        "leaf_senesced_cu_per_dry_mass",
        "leaf_senesced_fe_per_dry_mass",
        "leaf_senesced_k_per_dry_mass",
        "leaf_senesced_mg_per_dry_mass",
        "leaf_senesced_mn_per_dry_mass",
        "leaf_senesced_na_per_dry_mass",
        "leaf_senesced_s_per_dry_mass",
        "leaf_senesced_zn_per_dry_mass",
        "leaf_posture_numeric",
        "fire_fuel_bed_bulk_density",
        "fire_fuel_consumption",
        "fire_rate_of_spread",
        "fruit_ca_per_dry_mass",
        "fruit_k_per_dry_mass",
        "fruit_mg_per_dry_mass",
        "fruit_n_per_dry_mass",
        "fruit_s_per_dry_mass",
        "seed_ca_per_seed_dry_mass",
        "seed_k_per_seed_dry_mass",
        "seed_mg_per_seed_dry_mass",
        "seed_n_per_seed_dry_mass",
        "seed_s_per_seed_dry_mass",
        "stem_n_per_dry_mass",
        "wood_ca_per_dry_mass",
        "wood_k_per_dry_mass",
        "wood_mg_per_dry_mass",
        "wood_na_per_dry_mass",
        "wood_dead_ca_per_dry_mass",
        "wood_dead_k_per_dry_mass",
        "wood_dead_mg_per_dry_mass",
        "wood_dead_n_per_dry_mass",
        "wood_dead_na_per_dry_mass",
        "wood_dead_p_per_dry_mass",
        "flower_n_per_dry_mass",
        "leaf_al_per_dry_mass",
        "leaf_fresh_mass_per_area",
        "leaf_photosynthetic_nitrogen_use_efficiency_maximum",
        "leaf_photosynthetic_phosphorus_use_efficiency_maximum",
        "leaf_n_resorption",
        "leaf_p_resorption",
        "leaf_cell_wall_n_per_cell_wall_dry_mass",
        "leaf_cell_wall_n_per_leaf_n",
        "leaf_cell_wall_fraction",
        "leaf_guard_cell_length",
        "leaf_senesced_mo_per_dry_mass",
        "leaf_epidermis_ca_per_fresh_mass",
        "leaf_epidermis_p_per_fresh_mass",
        "leaf_hypodermis_ca_per_fresh_mass",
        "leaf_hypodermis_p_per_fresh_mass",
        "leaf_palisade_mesophyll_ca_per_fresh_mass",
        "leaf_palisade_mesophyll_p_per_fresh_mass",
        "leaf_sclerenchyma_ca_per_fresh_mass",
        "leaf_sclerenchyma_p_per_fresh_mass",
        "leaf_spongy_mesophyll_ca_per_fresh_mass",
        "leaf_spongy_mesophyll_p_per_fresh_mass",
        "leaf_internal_parenchyma_ca_per_fresh_mass",
        "leaf_internal_parenchyma_p_per_fresh_mass",
        "accessory_cost_fraction",
        "accessory_cost_mass",
        "leaf_cl_per_dry_mass",
        "leaf_photosynthesis_jmax_over_vcmax_25c",
        "leaf_photosynthesis_vcmax_per_area_25c",
        "ploidy",
        "leaf_epidermis_thickness",
        "leaf_palisade_cell_width",
        "leaf_stomatal_hairs",
        "leaf_total_vein_density",
        "leaf_cr_per_dry_mass",
        "leaf_co_per_dry_mass",
        "leaf_se_per_dry_mass",
        "seed_protein_per_seed_dry_mass",
        "plant_type_by_resource_use",
        "plant_tolerance_salt",
        "plant_tolerance_soil_salinity",
        "seed_oil_per_seed_dry_mass",
        "bark_dry_mass_per_surface_area",
        "wood_axial_parenchyma_fraction",
        "wood_conduit_fraction",
        "wood_ray_parenchyma_fraction",
        "stem_dry_matter_content",
        "stem_c_per_dry_mass",
        "leaf_mass_to_stem_mass_ratio",
        "diaspore_dry_mass",
        "root_c_per_dry_mass",
        "root_n_per_dry_mass",
        "root_delta13c",
        "root_delta15n",
        "leaf_epidermal_cell_density_abaxial",
        "leaf_epidermal_cell_density_adaxial",
        "stem_vessel_non_lumen_fraction",
        "leaf_inorganic_p_per_dry_mass",
        "leaf_lipid_p_per_dry_mass",
        "leaf_metabolite_p_per_dry_mass",
        "leaf_nucleic_acid_p_per_dry_mass",
        "leaf_residual_p_per_dry_mass",
        "leaf_lamina_mass_per_area",
        "plant_physical_defence_structures",
        "root_diameter",
        "root_distribution_coefficient",
        "root_dry_matter_content",
        "root_specific_root_area",
        "root_fine_root_coarse_root_ratio",
        "leaf_hairs_immature_leaves",
        "resprouting_capacity_stem_ratio",
        "flower_count_maximum",
        "flower_colour",
        "leaf_mesophyll_conductance_per_area",
        "leaf_mesophyll_conductance_per_mass",
        "bark_modulus_of_elasticity",
        "modulus_of_rupture",
        "stem_modulus_of_elasticity",
        "xylem_modulus_of_elasticity",
        "leaf_starch_per_area",
        "leaf_gas_exchange_quantum_yield",
        "stem_vessel_wall_fraction",
        "fruit_wall_thickness",
        "leaf_chlorophyll_per_dry_mass",
        "leaf_vessel_wall_thickness",
        "leaflet_count",
        "seed_colour",
        "leaf_work_to_tear",
        "stem_water_content_per_saturated_mass",
        "leaf_chlorophyll_index_modified_nd705",
        "bark_photosynthetic_status",
        "plant_height_reproductive",
        "storage_organ_diameter",
        "storage_organ_length",
        "stem_length",
        "flower_diameter",
        "resprouting_capacity_non_fire_disturbance",
        "plant_alternative_energy_and_nutrient_acquisition_strategy",
        "foliage_time",
        "flowering_cues",
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        "stem_delta13c",
        "stem_delta15n",
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        "leaf_chlorophyll_a_per_dry_mass",
        "leaf_chlorophyll_b_per_dry_mass",
        "leaf_epidermal_cell_density_both_sides",
        "leaf_insoluble_protein_per_area",
        "leaf_palisade_cell_length",
        "leaf_palisade_layer_number",
        "leaf_xylem_delta15n",
        "root_xylem_delta15n",
        "bud_length",
        "bud_width",
        "buds_per_inflorescence",
        "leaf_k_per_area",
        "root_specific_taproot_length",
        "stem_count",
        "sprout_depth",
        "leaf_secondary_vein_angle",
        "leaf_vein_frequency",
        "leaf_delta18o",
        "leaf_carotenoid_per_dry_mass",
        "leaf_rubisco_n_per_total_leaf_n",
        "leaf_rubisco_per_leaf_dry_mass",
        "leaf_thylakoid_n_per_total_leaf_n",
        "leaf_total_non_structural_carbohydrates_per_area",
        "leaf_total_non_structural_carbohydrates_per_mass",
        "reproductive_light_environment_index",
        "establishment_light_environment_index",
        "cotyledon_function",
        "cotyledon_position",
        "seedling_germination_location",
        "root_p_per_dry_mass",
        "branch_terminal_twig_cross_sectional_area",
        "seed_dry_mass_reserve",
        "branch_terminal_twig_length",
        "leaf_vessel_density",
        "plant_tolerance_snow",
        "plant_tolerance_water_logged_soils",
        "plant_tolerance_inundation",
        "plant_tolerance_calcicole",
        "bark_morphology_eucalyptus",
        "plant_spinescence",
        "seed_volume",
        "cotyledon_hairs",
        "embryo_colour",
        "seedling_first_node_leaf_count",
        "seedling_first_node_leaf_type",
        "seedling_hypocotyl_hairs",
        "bark_water_content_per_saturated_mass",
        "leaf_carotenoid_per_area",
        "leaf_chlorophyll_a_per_area",
        "leaf_chlorophyll_b_per_area",
        "leaf_absorption",
        "leaf_transmission",
        "bark_delta13c",
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        "type": "misc",
        "authors": "WoRMS Editorial Board",
        "year": "2026",
        "title": "World Register of Marine Species",
        "url": "https://www.marinespecies.org",
        "doi": "10.14284/170"
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      "license": "CC BY 4.0",
      "source_doi": "10.14284/170",
      "group_col": "region_code"
    }
  }
}
